The Paleozoic or Palaeozoic Era (from the Greek palaios (παλαιός), "old" and zoe (ζωή), "life", meaning "ancient life") is the earliest of three geologic Geology is the science and study of the physical matter and energy that constitute the Earth. The field of geology encompasses the study of the composition, structure, properties, and history of the planet's physical material, the processes by which it is formed, moved, and changed, the history of life on Earth, and human interactions with the eras The geologic time scale provides a system of chronologic measurement relating stratigraphy to time that is used by geologists, paleontologists and other earth scientists to describe the timing and relationships between events that have occurred during the history of the Earth. The table of geologic time spans presented here agrees with the dates of the Phanerozoic The Phanerozoic Eon is the current eon in the geologic timescale, and the one during which abundant animal life has existed. It covers roughly 545 million years and goes back to the time when diverse hard-shelled animals first appeared. Its name is derived from the Greek words φαίνω and ζωή, meaning make life appear, since it was once Eon The geologic time scale provides a system of chronologic measurement relating stratigraphy to time that is used by geologists, paleontologists and other earth scientists to describe the timing and relationships between events that have occurred during the history of the Earth. The table of geologic time spans presented here agrees with the dates. The Paleozoic spanned from roughly 542 to 251 million years ago (ICS, 2004), and is subdivided into six geologic periods The geologic time scale provides a system of chronologic measurement relating stratigraphy to time that is used by geologists, paleontologists and other earth scientists to describe the timing and relationships between events that have occurred during the history of the Earth. The table of geologic time spans presented here agrees with the dates; from oldest to youngest they are: the Cambrian The Cambrian is the first geological period of the Paleozoic Era, lasting from 542 ± 0.3 million years ago to 488.3 ± 1.7 million years ago(ICS, 2004,; it is succeeded by the Ordovician. Its subdivisions, and indeed its base, are somewhat in flux. The period was established by Adam Sedgwick, who named it after Cambria, the classical name for, Ordovician The Ordovician [/ɔɹdəˈvɪʃən/] is a geologic period and system, the second of six of the Paleozoic Era, and covers the time between 488.3±1.7 to 443.7±1.5 million years ago (ICS, 2004,. It follows the Cambrian Period and is followed by the Silurian Period. The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles, Silurian The Silurian is a geologic period and system that extends from the end of the Ordovician Period, about 443.7 ± 1.5 Ma , to the beginning of the Devonian Period, about 416.0 ± 2.8 Ma (ICS, 2004,. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by several, Devonian The Devonian is a geologic period and system of the Paleozoic Era spanning from 416 to 359.2 million years ago (ICS, 2004,. It is named after Devon, England, where rocks from this period were first studied, Carboniferous The Carboniferous is a geologic period and system that extends from the end of the Devonian Period, about 359.2 ± 2.5 Ma , to the beginning of the Permian Period, about 299.0 ± 0.8 Ma (ICS, 2004,, and Permian The Permian[note 1] is a geologic period and system characterized among land vertebrates by the diversification of the early amniotes into the ancestral groups of the mammals, turtles, lepidosaurs and archosaurs. The Permian Period follows the Carboniferous and extends from 299.0 ± 0.8 to 251.0 ± 0.4 Ma . It is the last period of the Paleozoic. Fish A fish is any aquatic vertebrate animal that is covered with scales, and equipped with two sets of paired fins and several unpaired fins. Most fish are "cold-blooded", or ectothermic, allowing their body temperatures to vary as ambient temperatures change. Fish are abundant in most bodies of water. They can be found in nearly all aquatic populations exploded in the Devonian. During the late Paleozoic, great forests A forest is an area with a high density of trees. There are many definitions of a forest, based on the various criteria. These plant communities cover approximately 9.4% of the Earth's surface (or 30% of total land area), though they once covered much more (about 50% of total land area), in many different regions and function as habitats for of primitive plants thrived on land forming the great coal Coal is a readily combustible black or brownish-black sedimentary rock normally occurring in rock strata in layers or veins called coal beds. The harder forms, such as anthracite coal, can be regarded as metamorphic rock because of later exposure to elevated temperature and pressure. Coal is composed primarily of carbon along with variable beds of Europe Europe is one of the world's seven continents. Comprising the westernmost peninsula of Eurasia, Europe is generally divided from Asia to its east by the water divide of the Ural Mountains, the Ural River, the Caspian Sea, the Caucasus region (Specification of borders) and the Black Sea to the southeast. Europe is bordered by the Arctic Ocean and and eastern North America. By the end of the era, the first large, sophisticated reptiles and the first modern plants (conifers The conifers, division Pinophyta, also known as division Coniferophyta or Coniferae, are one of 13 or 14 division level taxa within the Kingdom Plantae. Pinophytes are gymnosperms. They are cone-bearing seed plants with vascular tissue; all extant conifers are woody plants, the great majority being trees with just a few being shrubs. Typical) had developed.

The Paleozoic Era ended with the largest mass extinction in Earth's history, the Permian-Triassic Extinction Event The Permian–Triassic extinction event, informally known as the Great Dying, was an extinction event that occurred 251.4 million years ago, forming the boundary between the Permian and Triassic geologic periods. It was the Earth's most severe extinction event, with up to 96 percent of all marine species and 70 percent of terrestrial vertebrate. The effects of this catastrophe were so devastating that it took life on land 30M years to recover.^[1]. There are now indications that the recovery may have been much faster ^[2]

Geology

During the hiatus between the late Precambrian and the Paleozoic most of the evidence of the earth's early history was destroyed by erosion. From the beginning of the Paleozoic, shallow seas began to encroach on the continents.

In North America North America is the northern continent of the Americas, situated in the Earth's northern hemisphere and in the western hemisphere. It is bordered on the north by the Arctic Ocean, on the east by the North Atlantic Ocean, on the southeast by the Caribbean Sea, and on the west by the North Pacific Ocean; South America lies to the southeast, the era began with submerged geosynclines, or downward thrusts of the earth's crust, along the eastern, southeastern, and western sides of the continent, while the interior was dry land. As the era proceeded, the marginal seas periodically washed over the stable interior, leaving sedimentary deposits to mark their incursions. During the early part of the era, the area of exposed Precambrian, or shield, rocks in central Canada The land occupied by Canada was inhabited for millennia by various groups of Aboriginal peoples. Beginning in the late 15th century, British and French expeditions explored, and later settled, along the Atlantic coast. France ceded nearly all of its colonies in North America in 1763 after the Seven Years' War. In 1867, with the union of three were eroding, supplying sediment to the geosynclines from the interior. Beginning in the Ordovician period, mountain building intermittently proceeded in the eastern part of the Appalachian geosyncline throughout the rest of the era, bringing in new sediments. Sediments washing from the Acadian Mountains filled the western part of the Appalachian geosyncline to form the famous coal Coal is a readily combustible black or brownish-black sedimentary rock normally occurring in rock strata in layers or veins called coal beds. The harder forms, such as anthracite coal, can be regarded as metamorphic rock because of later exposure to elevated temperature and pressure. Coal is composed primarily of carbon along with variable swamps of the Carboniferous period. Uplift of the Appalachians meant that the region was never again inundated by vast marginal seas.

Paleoclimatic studies and evidence of glaciers indicate that central Africa was most likely in the polar regions during the early Paleozoic. During the early Paleozoic, the huge continent Gondwanaland had either formed or was forming. By mid-Paleozoic, the collision of North America and Europe produced the Acadian-Caledonian uplifts, and a subduction plate uplifted eastern Australia. By the late Paleozoic, continental collisions formed the supercontinent Pangaea and resulted in some of the great mountain chains, including the Appalachians, Urals, and Tasmans.

Life

The most noteworthy feature of Paleozoic life is the sudden appearance of nearly all of the invertebrate An invertebrate is an animal without a backbone. The group includes 95% of all animal species — all animals except those in the chordate subphylum Vertebrata animal phyla in great abundance at the beginning of the Cambrian. A few primitive fishlike invertebrates, and then vertebrates Vertebrates are members of the subphylum Vertebrata, chordates with backbones and spinal columns. About 58,000 species of vertebrates have been currently described. Vertebrata is the largest subphylum of chordates, and contains many familiar groups of large land animals. Vertebrates are comprised of animals from the groups jawless fishs, bony fish,, appeared in the Cambrian and Ordovician, scorpions Scorpions are predatory arthropod animals of the order Scorpiones within the class Arachnida. They have eight legs and are easily recognised by the pair of grasping claws and the narrow, segmented tail, carried in a characteristic forward curve over the back, ending with a venomous stinger. Though the scorpion has a fearsome reputation as venomous, in the Silurian period, land invertebrates and amphibians Amphibians , such as frogs, toads, salamanders, newts, and caecilians, are ectothermic (or cold-blooded) animals that metamorphose from a juvenile water-breathing form, either to an adult air-breathing form, or to a paedomorph that retains some juvenile characteristics. Proteidae (mudpuppies and waterdogs) are good examples of paedomorphic species in the Devonian, land reptiles Reptiles are animals in the class Reptilia characterized by breathing air, a "cold-blooded" (poikilothermic) metabolism, laying tough-shelled amniotic eggs (or retaining the same membrane system in species with live birth), and skin with scales or scutes. They are tetrapods (either having four limbs or being descended from four-limbed in the Carboniferous, and marine reptiles in the Permian. All reptiles increased in number and in variety by the late Permian. The plant life of the Paleozoic era reached its climax in the Carboniferous, and was then much diminished in the Permian.

Tectonics

Geologically, the Paleozoic starts shortly after the breakup of a supercontinent In geology, a supercontinent is a landmass comprising more than one continental core, or craton. The assembly of cratons and accreted terranes that form Eurasia qualifies as a supercontinent today called Pannotia Pannotia, first described by Ian W. D. Dalziel in 1997, is a hypothetical supercontinent that existed from the Pan-African orogeny about 600 million years ago to the end of the Precambrian about 540 million years ago. It is also known as the Vendian supercontinent and at the end of a global ice age An "ice age" or, more precisely, "glacial age" is a generic geological period of long-term reduction in the temperature of the Earth's surface and atmosphere, resulting in an expansion of continental ice sheets, polar ice sheets and alpine glaciers. Within a long-term ice age, individual pulses of extra cold climate are termed &. (See Varanger glaciation The Cryogenian is a geologic period that lasted from 850 to 635 million years ago. The Cryogenian forms the second geologic period of the Neoproterozoic Era, preceded by the Tonian period and followed by the Ediacaran. The Sturtian and Marinoan glaciations, which are the greatest ice ages known to have occurred on Earth and may have covered the and Snowball Earth Snowball Earth refers to the hypothesis that the Earth's surface became nearly or entirely frozen at least once, before 650 million years ago. The geological community generally accepts this hypothesis because it best explains sedimentary deposits generally regarded as of glacial origin at tropical paleolatitudes and other enigmatic features in). Throughout the early Paleozoic, the Earth's landmass was broken up into a substantial number of relatively small continents. Toward the end of the era, the continents gathered together into a supercontinent In geology, a supercontinent is a landmass comprising more than one continental core, or craton. The assembly of cratons and accreted terranes that form Eurasia qualifies as a supercontinent today called Pangaea Pangaea, Pangæa, or Pangea was the supercontinent that existed during the Paleozoic and Mesozoic eras about 250 million years ago, before the component continents were separated into their current configuration, which included most of the Earth's land area.

Climate

The Early Cambrian climate was probably moderate at first, becoming warmer over the course of the Cambrian, as the second-greatest sustained sea level rise Current sea level rise has occurred at a mean rate of 1.8 mm per year for the past century, and more recently, during the satellite era of sea level measurement, at rates estimated near 2.8 ± 0.4 to 3.1 ± 0.7 mm per year . Current sea level rise is due significantly to global warming, which will increase sea level over the coming century and in the Phanerozoic got underway. However, as if to offset this trend, Gondwana Gondwana , originally Gondwanaland, is the name given to a southern precursor supercontinent. Its final geological suturing occurred between ca. 570 and 510 million years ago (Ma), joining East Gondwana to West Gondwana. It later separated from Laurasia 180-200 million years ago during the breakup of the Pangaea supercontinent that existed about 50 moved south with considerable speed, so that, in Ordovician time, most of West Gondwana (Africa and South America) lay directly over the South Pole The South Pole, also known as the Geographic South Pole or Terrestrial South Pole, is one of the two points where the Earth's axis of rotation intersects its surface. It is the southernmost point on the surface of the Earth and lies on the opposite side of the Earth from the North Pole. Situated on the continent of Antarctica, it is the site of. The Early Paleozoic climate was also strongly zonal, with the result that the "climate", in an abstract sense became warmer, but the living space of most organisms of the time—the continental shelf marine environment—became steadily colder. However, Baltica Baltica is a name applied by geologists to a late-Proterozoic, early-Palaeozoic continent that now includes the East European craton of northwestern Eurasia. Baltica was created as an entity not earlier than 1.8 billion years ago. Before this time, the three segments/continents that now comprise the East European craton were in different places on (Northern Europe and Russia) and Laurentia Laurentia , like all craton land, was created as continents moved about the surface of the Earth, bumping into other continents and drifting away (eastern North America and Greenland) remained in the tropical zone, while China and Australia lay in waters which were at least temperate. The Early Paleozoic ended, rather abruptly, with the short, but apparently severe, Late Ordovician Ice Age. This cold spell caused the second-greatest mass extinction of Phanerozoic time. Over time, the warmer weather moved into the Paleozoic era.

The Middle Paleozoic was a time of considerable stability. Sea levels had dropped coincident with the Ice Age, but slowly recovered over the course of the Silurian and Devonian. The slow merger of Baltica and Laurentia, and the northward movement of bits and pieces of Gondwana created numerous new regions of relatively warm, shallow sea floor. As plants took hold on the continental margins, oxygen Oxygen (pronounced /ˈɒksɨdʒɨn/, OK-si-jin, from the Greek roots ὀξύς (acid, literally "sharp", from the taste of acids) and -γενής (-genēs) (producer, literally begetter), is the element with atomic number 8 and represented by the symbol O. It is a member of the chalcogen group on the periodic table, and is a highly levels increased and carbon dioxide Carbon dioxide is a chemical compound composed of two oxygen atoms covalently bonded to a single carbon atom. It is a gas at standard temperature and pressure and exists in Earth's atmosphere in this state. CO2 is a trace gas comprising 0.039% of the atmosphere dropped, although much less dramatically. The north-south temperature gradient also seems to have moderated, or metazoan life simply became hardier, or both. At any event, the far southern continental margins of Antarctica and West Gondwana became increasingly less barren. The Devonian ended with a series of turnover pulses which killed off much of Middle Paleozoic vertebrate life, without noticeably reducing species diversity overall.

The Late Paleozoic was a time which has left us a good many unanswered questions. The Mississippian epoch The Mississippian is a subperiod in the geologic timescale or a subsystem of the geologic record. It is the earliest/lowermost of two subperiods of the Carboniferous period lasting from roughly 359 to 318 Ma . As with most other geochronologic units, the rock beds that define the Mississippian are well identified, but the exact start and end dates began with a spike in atmospheric oxygen, while carbon dioxide plummeted to unheard-of lows. This destabilized the climate and led to one, and perhaps two, ice ages during the Carboniferous The Carboniferous is a geologic period and system that extends from the end of the Devonian Period, about 359.2 ± 2.5 Ma , to the beginning of the Permian Period, about 299.0 ± 0.8 Ma (ICS, 2004,. These were far more severe than the brief Late Ordovician Ice; but, this time, the effects on world biota were inconsequential. By the Cisuralian, both oxygen and carbon dioxide had recovered to more normal levels. On the other hand, the assembly of Pangea created huge arid inland areas subject to temperature extremes. The Lopingian The Permian[note 1] is a geologic period and system characterized among land vertebrates by the diversification of the early amniotes into the ancestral groups of the mammals, turtles, lepidosaurs and archosaurs. The Permian Period follows the Carboniferous and extends from 299.0 ± 0.8 to 251.0 ± 0.4 Ma . It is the last period of the Paleozoic is associated with falling sea levels, increased carbon dioxide and general climatic deterioration, culminating in the devastation of the Permian extinction.

See also

• Geologic timescale The geologic time scale is a chronologic schema relating stratigraphy to time that is used by geologists, paleontologists and other earth scientists to describe the timing and relationships between events that have occurred during the history of the Earth. The table of geologic time spans presented here agrees with the dates and nomenclature

References and further reading

1. ^ Sahney, S. and Benton, M.J. (2008). "Recovery from the most profound mass extinction of all time" (PDF). Proceedings of the Royal Society: Biological 275 (1636): 759. doi A digital object identifier is a character string used to uniquely identify an electronic document or other object. Metadata about the object is stored in association with the DOI name and this metadata may include a location, such as a URL, where the object can be found. The DOI for a document is permanent, whereas its location and other metadata:10.1098/rspb.2007.1370. PMID A PMID is a unique number assigned to each PubMed citation of life sciences and biomedical scientific journal articles. The related Pubmed Central archive may additionally assign a separate number, a PMCID (PubMed Central Identifier), normally written with a PMC prefix 18198148. PMC PubMed Central is a free digital database of full-text scientific literature in biomedical and life sciences. It grew from the online Entrez PubMed biomedical literature search system. PubMed Central was developed by the U.S. National Library of Medicine as an online archive of biomedical journal articles 2596898. http://journals.royalsociety.org/content/qq5un1810k7605h5/fulltext.pdf.
2. ^ url=http://www.economist.com/node/16524904

• British Palaeozoic Fossils, 1975, The Natural History Museum, London.
• "International Commission on Stratigraphy (ICS)". Home Page. http://www.stratigraphy.org/. Retrieved September 19, 2005.

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