Plants are living Life is a characteristic that distinguishes objects that have signaling and self-sustaining processes (biology) from those that do not, either because such functions have ceased (death), or else because they lack such functions and are classified as inanimate organisms In biology, an organism is any contiguous living system . In at least some form, all organisms are capable of response to stimuli, reproduction, growth and development, and maintenance of homoeostasis as a stable whole. An organism may either be unicellular (single-celled) or be composed of, as in humans, many trillions of cells grouped into belonging to the kingdom In biology, kingdom or regnum is a taxonomic rank, which is either the highest rank or in the more recent three-domain system, the rank below domain. Kingdoms are divided into smaller groups called phyla or divisions in botany. The complete sequence of ranks is life, domain, kingdom, phylum, class, order, family, genus, and species Plantae. They include familiar organisms such as trees A tree is a perennial woody plant. It is most often defined as a woody plant that has many secondary branches supported clear of the ground on a single main stem or trunk with clear apical dominance. A minimum height specification at maturity is cited by some authors, varying from 3 m to 6 m; some authors set a minimum of 10 cm trunk diameter, herbs In American English the initial "h" is normally silent: /ˈɜrb/.[Full citation needed] In standard British English the "h" is pronounced: /ˈhɜːb/ Also see American and British English pronunciation differences. In Canada, it is pronounced either with or without the "h", bushes A shrub or bush is distinguished from a tree by its multiple stems and lower height, usually less than 5–6 m tall. A large number of plants can be either shrubs or trees, depending on the growing conditions they experience. Small, low shrubs such as lavender, periwinkle and thyme are often termed subshrubs, grasses Grasses, or more technically graminoids, are monocotyledonous, usually herbaceous plants with narrow leaves growing from the base. They include the "true grasses", of the Poaceae family, as well as the sedges (Cyperaceae) and the rushes (Juncaceae). The true grasses include cereals, bamboo and the grasses of lawns (turf) and grassland, vines A vine in the broad sense refers to any climbing or trailing plant. The narrower and original meaning is the grapevine (Vitis), ferns A fern is any one of a group of about 12,000 species of plants. Unlike mosses, they have xylem and phloem . They have stems, leaves, and roots like other vascular plants. Ferns do not have either seeds or flowers (they reproduce via spores), mosses Mosses are small, soft plants that are typically 1–10 cm tall, though some species are much larger. They commonly grow close together in clumps or mats in damp or shady locations. They do not have flowers or seeds, and their simple leaves cover the thin wiry stems. At certain times mosses produce spore capsules which may appear as beak-like, and green algae The green algae are the large group of algae from which the embryophytes (higher plants) emerged. As such, they form a paraphyletic group, although the group including both green algae and embryophytes is monophyletic (and often just known as kingdom Plantae). The green algae include unicellular and colonial flagellates, usually but not always. The scientific study of plants, known as botany Botany, plant science, phytology, or plant biology is a branch of biology that involves the scientific study of plant life. Botany covers a wide range of scientific disciplines concerned with the study of plants, algae and fungi, including structure, growth, reproduction, metabolism, development, diseases, chemical properties, and evolutionary, has identified about 350,000 extant Extant is a term commonly used in biology to refer to taxa that are still in existence (living). The term extant contrasts with extinct. For example, Brandt's Cormorant is an extant species, while the Spectacled Cormorant is an extinct species. Likewise, of the group of molluscs known as the cephalopods, there are approximately 600 extant species species In biology, a species is one of the basic units of biological classification and a taxonomic rank. A species is often defined as a group of organisms capable of interbreeding and producing fertile offspring. While in many cases this definition is adequate, more precise or differing measures are often used, such as based on similarity of DNA or of plants, defined as seed plants The spermatophytes (also known as phanerogams) comprise those plants that produce seeds. They are a subset of the embryophytes or land plants. The living spermatophytes form five groups:, bryophytes Bryophytes are all embryophytes that are non-vascular: they have tissues and enclosed reproductive systems, but they lack vascular tissue that circulates liquids. They neither have flowers nor produce seeds, reproducing via spores. The term bryophyte comes from Greek βρύον - bryon, "tree-moss, oyster-green" + φυτόν - fyton &, ferns A fern is any one of a group of about 12,000 species of plants. Unlike mosses, they have xylem and phloem . They have stems, leaves, and roots like other vascular plants. Ferns do not have either seeds or flowers (they reproduce via spores) and fern allies Fern ally is a general term covering a somewhat diverse group of seedless vascular plants that are not true ferns. Like ferns, these plants disperse by shedding spores to initiate an alternation of generations. As of 2004, some 287,655 species had been identified, of which 258,650 are flowering A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants . The biological function of a flower is to mediate the union of male sperm with female ovum in order to produce seeds. The process begins with pollination, is followed by fertilization, leading to the formation and dispersal of the seeds. For and 18,000 bryophytes Bryophytes are all embryophytes that are non-vascular: they have tissues and enclosed reproductive systems, but they lack vascular tissue that circulates liquids. They neither have flowers nor produce seeds, reproducing via spores. The term bryophyte comes from Greek βρύον - bryon, "tree-moss, oyster-green" + φυτόν - fyton & (see table below). Green plants, sometimes called Viridiplantae, obtain most of their energy from sunlight Electromagnetic radiation is a phenomenon that takes the form of self-propagating waves in a vacuum or in matter. It comprises electric and magnetic field components, which oscillate in phase perpendicular to each other and perpendicular to the direction of energy propagation. Electromagnetic radiation is classified into several types according to via a process called photosynthesis Photosynthesis is a process that converts carbon dioxide into organic compounds, especially sugars, using the energy from sunlight. Photosynthesis occurs in plants, algae, and many species of bacteria, but not in archaea. Photosynthetic organisms are called photoautotrophs, since they can create their own food. In plants, algae, and cyanobacteria,.

Contents

Definition

Aristotle Aristotle (384 BC – 322 BC) was a Greek philosopher, a student of Plato and teacher of Alexander the Great. His writings cover many subjects, including physics, metaphysics, poetry, theater, music, logic, rhetoric, politics, government, ethics, biology, and zoology. Together with Plato and Socrates (Plato's teacher), Aristotle is one of the most divided all living things between plants (which generally do not move), and animals (which often are mobile to catch their food). In Linnaeus Carl Linnaeus [a 2] (Latinized as Carolus Linnaeus [a 3], also known after his ennoblement as Carl von Linné , 23 May[a 1] 1707 – 10 January 1778) was a Swedish botanist, physician, and zoologist, who laid the foundations for the modern scheme of binomial nomenclature. He is known as the father of modern taxonomy, and is also considered one of' system, these became the Kingdoms In biology, kingdom or regnum is a taxonomic rank, which is either the highest rank or in the more recent three-domain system, the rank below domain. Kingdoms are divided into smaller groups called phyla or divisions in botany. The complete sequence of ranks is life, domain, kingdom, phylum, class, order, family, genus, and species Vegetabilia (later Metaphyta or Plantae) and Animalia Animals are a major group of mostly multicellular, eukaryotic organisms of the kingdom Animalia or Metazoa. Their body plan eventually becomes fixed as they develop, although some undergo a process of metamorphosis later on in their life. Most animals are motile, meaning they can move spontaneously and independently. All animals are also (also called Metazoa). Since then, it has become clear that the Plantae as originally defined included several unrelated groups, and the fungi A fungus is a member of a large group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. The Fungi (pronounced /ˈfʌndʒaɪ/ or /ˈfʌŋɡaɪ/) are classified as a kingdom that is separate from plants, animals and bacteria. One major difference is that fungal cells have cell and several groups of algae Algae are a large and diverse group of simple, typically autotrophic organisms, ranging from unicellular to multicellular forms. The largest and most complex marine forms are called seaweeds. They are photosynthetic, like plants, and "simple" because they lack the many distinct organs found in land plants were removed to new kingdoms. However, these are still often considered plants in many contexts, both technical and popular.

Current definitions of Plantae

When the name Plantae or plants is applied to a specific taxon, it is usually referring to one of three concepts. From smallest to largest in inclusiveness, these three groupings are:

Name(s) Scope Description
Land plants The embryophytes are the most familiar group of plants. They are often called land plants because they live primarily in terrestrial habitats, in contrast with the related green algae that are primarily aquatic. The embryophytes include trees, flowers, ferns, mosses, and various other green land plants. All are complex multicellular eukaryotes, also known as Embryophyta or Metaphyta. Plantae sensu strictissimo In rare cases the superlatives sensu strictissimo and sensu latissimo ("in the widest sense") may be used. Another common usage is in conjunction with an author citation, indicating that the intended meaning is the one defined by that author As the narrowest of plant categories, this is further delineated below.
Green plants - also known as Viridiplantae Viridiplantae are a clade comprising the green algae and land plants, Viridiphyta or Chlorobionta Plantae sensu stricto In rare cases the superlatives sensu strictissimo and sensu latissimo ("in the widest sense") may be used. Another common usage is in conjunction with an author citation, indicating that the intended meaning is the one defined by that author Comprise the above Embryophytes, Charophyta The Charophyta are a division of green algae, including the closest relatives of the embryophyte plants. In some groups, such as conjugating green algae, flagellate cells do not occur. The latter group does engage in sexual reproduction, and motility does not involve flagella, since they are totally lacking. Flagellate cells in the form of sperm (i.e., primitive stoneworts Charales is an order of pondweeds, freshwater algae in the division Charophyta. They are green plants believed to be the closest relatives of the green land plants. Linnaeus established the genus in 1753), and Chlorophyta Chlorophyta, a division of green algae, includes about 7,000 species of mostly aquatic photosynthetic eukaryotic organisms. Like the land plants , green algae contain chlorophylls a and b, and store food as starch in their plastids. They are related to the Charophyta and Embryophyta (land plants), together making up the Viridiplantae (i.e., green algae The green algae are the large group of algae from which the embryophytes (higher plants) emerged. As such, they form a paraphyletic group, although the group including both green algae and embryophytes is monophyletic (and often just known as kingdom Plantae). The green algae include unicellular and colonial flagellates, usually but not always such as sea lettuce The sea lettuces comprise the genus Ulva, a group of edible green algae that are widely distributed along the coasts of the world's oceans. The type species within the genus Ulva is Ulva lactuca Linnaeus, "lactuca" meaning lettuce. Additionally, some species in the algae genus Monostroma are known as slender sea lettuces). Viridiplantae encompasses a group of organisms that possess chlorophyll Chlorophyll is a green pigment found in all plants, algae, and cyanobacteria. Its name is derived from the Greek χλωρός and φύλλον (phyllon "leaf"). Chlorophyll absorbs light most strongly in the blue portion of the electromagnetic spectrum, followed by the red portion. However, it is a poor absorber of green and near-green a and b, have plastids Plastids are major organelles found in the cells of plants and algae. Plastids are the site of manufacture and storage of important chemical compounds used by the cell. Plastids often contain pigments used in photosynthesis, and the types of pigments present can change or determine the cell's colour that are bound by only two membranes, are capable of storing starch, and have cellulose Cellulose is an organic compound with the formula n, a polysaccharide consisting of a linear chain of several hundred to over ten thousand β(1→4) linked D-glucose units in their cell walls A cell wall is a tough, usually flexible but sometimes fairly rigid layer that surrounds some types of cells. It is located outside the cell membrane and provides these cells with structural support and protection, and also acts as a filtering mechanism. A major function of the cell wall is to act as a pressure vessel, preventing over-expansion. It is this clade A clade[note 1] is a group consisting of an organism and all its descendants. In the terms of biological systematics, a clade is a single "branch" on the "tree of life". The idea that such a "natural group" of organisms should be grouped together and given a taxonomic name is central to biological classification. In which is mainly the subject of this article.
Archaeplastida The Archaeplastida are a major group of eukaryotes, comprising the red and green algae and the land plants, together with a small group called the glaucophytes. The plastids of all of these organisms are surrounded by two membranes, suggesting they developed directly from endosymbiotic cyanobacteria. In all other groups, plastids are surrounded by, Plastida or Primoplantae Plantae sensu lato In rare cases the superlatives sensu strictissimo and sensu latissimo ("in the widest sense") may be used. Another common usage is in conjunction with an author citation, indicating that the intended meaning is the one defined by that author Comprises the green plants above, as well as Rhodophyta The Red algae (or Rhodophyta, pronounced /roʊˈdɒfɨtə, ˌroʊdəˈfaɪtə/, from Greek: ῥόδον = rose + φυτόν (phyton) = plant, thus red plant) are one of the oldest groups of eukaryotic algae, and also one of the largest, with about 5,000–6,000 species of mostly multicellular, marine algae, including many notable seaweeds. Other (red algae) and Glaucophyta The glaucophytes, also known as glaucocystophytes or glaucocystids, are a small group of freshwater microscopic algae. Together with the red algae and green algae plus land plants (Viridiplantae or Chloroplastida), they form the Archaeplastida. However, the relationships between the red algae, green algae and glaucophytes are unclear, in large (simple glaucophyte algae). As the broadest plant clade, this comprises most of the eukaryotes A eukaryote is an organism whose cells contain complex structures enclosed within membranes. The defining membrane-bound structure that sets eukaryotic cells apart from prokaryotic cells is the nucleus, or nuclear envelope, within which the genetic material is carried. The presence of a nucleus gives eukaryotes their name, which comes from the that eons ago acquired their chloroplasts Chloroplasts are organelles found in plant cells and other eukaryotic organisms that conduct photosynthesis. Chloroplasts capture light energy to conserve free energy in the form of ATP and reduce NADP to NADPH through a complex set of processes called photosynthesis directly by engulfing cyanobacteria Cyanobacteria is a phylum of bacteria that obtain their energy through photosynthesis. The name "cyanobacteria" comes from the color of the bacteria (Greek: κυανός (kyanós) = blue).

Outside of formal scientific contexts, the term "plant" implies an association with certain traits, such as multicellularity, cellulose, and photosynthesis.[2][3] Many of the classification controversies involve organisms that are rarely encountered and are of minimal apparent economic significance, but are crucial in developing an understanding of the evolution of modern flora.

Algae

Green algae The green algae are the large group of algae from which the embryophytes (higher plants) emerged. As such, they form a paraphyletic group, although the group including both green algae and embryophytes is monophyletic (and often just known as kingdom Plantae). The green algae include unicellular and colonial flagellates, usually but not always from Ernst Haeckel's Kunstformen der Natur, 1904. Main article: Algae

Most algae are no longer classified within the Kingdom Plantae.[4][5] The algae comprise several different groups of organisms that produce energy through photosynthesis, each of which arose independently from separate non-photosynthetic ancestors. Most conspicuous among the algae are the seaweeds, multicellular algae that may roughly resemble terrestrial plants, but are classified among the green, red, and brown algae. Each of these algal groups also includes various microscopic and single-celled organisms.

The two groups of green algae are the closest relatives of land plants (embryophytes). The first of these groups is the Charophyta (desmids and stoneworts), from which the embryophytes developed.[6][7][8] The sister group to the combined embryophytes and charophytes is the other group of green algae, Chlorophyta, and this more inclusive group is collectively referred to as the green plants or Viridiplantae. The Kingdom Plantae is often taken to mean this monophyletic grouping. With a few exceptions among the green algae, all such forms have cell walls containing cellulose, have chloroplasts containing chlorophylls a and b, and store food in the form of starch. They undergo closed mitosis without centrioles, and typically have mitochondria with flat cristae.

The chloroplasts of green plants are surrounded by two membranes, suggesting they originated directly from endosymbiotic cyanobacteria. The same is true of two additional groups of algae: the Rhodophyta (red algae) and Glaucophyta. All three groups together are generally believed to have a common origin, and so are classified together in the taxon Archaeplastida. In contrast, most other algae (e.g. heterokonts, haptophytes, dinoflagellates, and euglenids) have chloroplasts with three or four surrounding membranes. They are not close relatives of the green plants, presumably acquiring chloroplasts separately from ingested or symbiotic green and red algae.

Fungi

Main article: Fungi

The classification of fungi has been controversial until quite recently in the history of biology. Linnaeus' original classification placed the fungi within the Plantae, since they were unquestionably not animalian; this being the only other alternative. With later developments in microbiology, in the 19th century Ernst Haeckel felt that a third kingdom was required to classify newly discovered micro-organisms. The introduction of the new kingdom Protista as an alternative to Animalia, led to uncertainty as to whether fungi truly were best placed in the Plantae or whether they ought to be reclassified as protists. Haeckel himself found it difficult to decide and it was not until 1969 that a solution was found whereby Robert Whittaker proposed the creation of the kingdom Fungi. Molecular evidence has since shown that the concestor (last common ancestor) of the Fungi was probably more similar to that of the Animalia than of any other kingdom, including the Plantae.

Whittaker's original reclassification was based on the fundamental difference in nutrition between the Fungi and the Plantae. Unlike plants, which are generally autotrophic multicellular phototrophs which gain carbon through photosynthesis, fungi are generally heterotrophic uni- or multi-cellular saprotrophs, obtaining carbon by breaking down and absorbing surrounding materials. In addition, the substructure of multicellular fungi takes the form of many chitinous microscopic strands called hyphae, which may be further subdivided into cells or may form a syncytium containing many eukaryotic nuclei. Fruiting bodies, of which mushrooms are most familiar example, are the reproductive structures of fungi.

Diversity

About 350,000 species of plants, defined as seed plants, bryophytes, ferns and fern allies, are estimated to exist currently. As of 2004, some 287,655 species had been identified, of which 258,650 are flowering plants, 16,000 bryophytes, 11,000 ferns and 8,000 green algae.

Diversity of living plant divisions
Informal group Division name Common name No. of living species
Green algae Chlorophyta green algae (chlorophytes) 3,800 [9]
Charophyta green algae (desmids & charophytes) 4,000 - 6,000 [10]
Bryophytes Marchantiophyta liverworts 6,000 - 8,000 [11]
Anthocerotophyta hornworts 100 - 200 [12]
Bryophyta mosses 12,000 [13]
Pteridophytes Lycopodiophyta club mosses 1,200 [5]
Pteridophyta ferns, whisk ferns & horsetails 11,000 [5]
Seed plants Cycadophyta cycads 160 [14]
Ginkgophyta ginkgo 1 [15]
Pinophyta conifers 630 [5]
Gnetophyta gnetophytes 70 [5]
Magnoliophyta flowering plants 258,650 [16]

The naming of plants is governed by the International Code of Botanical Nomenclature and International Code of Nomenclature for Cultivated Plants (see cultivated plant taxonomy).

Phylogeny

This article may be confusing or unclear to readers. Please help clarify the article; suggestions may be found on the talk page. (March 2009)

A proposed phylogenetic tree of Plantae, after Kenrick and Crane,[17] is as follows, with modification to the Pteridophyta from Smith et al.[18] The Prasinophyceae may be a paraphyletic basal group to all green plants.

Prasinophyceae (micromonads)

Streptobionta
Embryophytes
Stomatophytes
Polysporangiates
Tracheophytes
Eutracheophytes
Euphyllophytina
Lignophytia

Spermatophytes (seed plants)

Progymnospermophyta

Pteridophyta

Pteridopsida (true ferns)

Marattiopsida

Equisetopsida (horsetails)

Psilotopsida (whisk ferns & adders'-tongues)

Cladoxylopsida

Lycophytina

Lycopodiophyta

Zosterophyllophyta

Rhyniophyta

Aglaophyton

Horneophytopsida

Bryophyta (mosses)

Anthocerotophyta (hornworts)

Marchantiophyta (liverworts)

Charophyta

Chlorophyta

Trebouxiophyceae (Pleurastrophyceae)

Chlorophyceae

Ulvophyceae

Embryophytes

Main article: Embryophyte Dicksonia antarctica, a species of tree fern

The plants that are likely most familiar to us are the multicellular land plants, called embryophytes. They include the vascular plants, plants with full systems of leaves, stems, and roots. They also include a few of their close relatives, often called bryophytes, of which mosses and liverworts are the most common.

All of these plants have eukaryotic cells with cell walls composed of cellulose, and most obtain their energy through photosynthesis, using light and carbon dioxide to synthesize food. About three hundred plant species do not photosynthesize but are parasites on other species of photosynthetic plants. Plants are distinguished from green algae, which represent a mode of photosynthetic life similar to the kind modern plants are believed to have evolved from, by having specialized reproductive organs protected by non-reproductive tissues.

Bryophytes first appeared during the early Paleozoic. They can only survive where moisture is available for significant periods, although some species are desiccation tolerant. Most species of bryophyte remain small throughout their life-cycle. This involves an alternation between two generations: a haploid stage, called the gametophyte, and a diploid stage, called the sporophyte. The sporophyte is short-lived and remains dependent on its parent gametophyte.

Vascular plants first appeared during the Silurian period, and by the Devonian had diversified and spread into many different land environments. They have a number of adaptations that allowed them to overcome the limitations of the bryophytes. These include a cuticle resistant to desiccation, and vascular tissues which transport water throughout the organism. In most the sporophyte acts as a separate individual, while the gametophyte remains small.

The first primitive seed plants, Pteridosperms (seed ferns) and Cordaites, both groups now extinct, appeared in the late Devonian and diversified through the Carboniferous, with further evolution through the Permian and Triassic periods. In these the gametophyte stage is completely reduced, and the sporophyte begins life inside an enclosure called a seed, which develops while on the parent plant, and with fertilisation by means of pollen grains. Whereas other vascular plants, such as ferns, reproduce by means of spores and so need moisture to develop, some seed plants can survive and reproduce in extremely arid conditions.

Early seed plants are referred to as gymnosperms (naked seeds), as the seed embryo is not enclosed in a protective structure at pollination, with the pollen landing directly on the embryo. Four surviving groups remain widespread now, particularly the conifers, which are dominant trees in several biomes. The angiosperms, comprising the flowering plants, were the last major group of plants to appear, emerging from within the gymnosperms during the Jurassic and diversifying rapidly during the Cretaceous. These differ in that the seed embryo (angiosperm) is enclosed, so the pollen has to grow a tube to penetrate the protective seed coat; they are the predominant group of flora in most biomes today.

Fossils

Main articles: Paleobotany and Evolutionary history of plants A petrified log in Petrified Forest National Park.

Plant fossils include roots, wood, leaves, seeds, fruit, pollen, spores, phytoliths, and amber (the fossilized resin produced by some plants). Fossil land plants are recorded in terrestrial, lacustrine, fluvial and nearshore marine sediments. Pollen, spores and algae (dinoflagellates and acritarchs) are used for dating sedimentary rock sequences. The remains of fossil plants are not as common as fossil animals, although plant fossils are locally abundant in many regions worldwide.

The earliest fossils clearly assignable to Kingdom Plantae are fossil green algae from the Cambrian. These fossils resemble calcified multicellular members of the Dasycladales. Earlier Precambrian fossils are known which resemble single-cell green algae, but definitive identity with that group of algae is uncertain.

The oldest known fossils of embryophytes date from the Ordovician, though such fossils are fragmentary. By the Silurian, fossils of whole plants are preserved, including the lycophyte Baragwanathia longifolia. From the Devonian, detailed fossils of rhyniophytes have been found. Early fossils of these ancient plants show the individual cells within the plant tissue. The Devonian period also saw the evolution of what many believe to be the first modern tree, Archaeopteris. This fern-like tree combined a woody trunk with the fronds of a fern, but produced no seeds.

The Coal measures are a major source of Paleozoic plant fossils, with many groups of plants in existence at this time. The spoil heaps of coal mines are the best places to collect; coal itself is the remains of fossilised plants, though structural detail of the plant fossils is rarely visible in coal. In the Fossil Forest at Victoria Park in Glasgow, Scotland, the stumps of Lepidodendron trees are found in their original growth positions.

The fossilized remains of conifer and angiosperm roots, stems and branches may be locally abundant in lake and inshore sedimentary rocks from the Mesozoic and Cenozoic eras. Sequoia and its allies, magnolia, oak, and palms are often found.

Petrified wood is common in some parts of the world, and is most frequently found in arid or desert areas where it is more readily exposed by erosion. Petrified wood is often heavily silicified (the organic material replaced by silicon dioxide), and the impregnated tissue is often preserved in fine detail. Such specimens may be cut and polished using lapidary equipment. Fossil forests of petrified wood have been found in all continents.

Fossils of seed ferns such as Glossopteris are widely distributed throughout several continents of the Southern Hemisphere, a fact that gave support to Alfred Wegener's early ideas regarding Continental drift theory.

Structure, growth, and development

Further information: Plant morphology

Most of the solid material in a plant is taken from the atmosphere. Through a process known as photosynthesis, most plants use the energy in sunlight to convert carbon dioxide from the atmosphere, plus water, into simple sugars. Parasitic plants, on the other hand, use the resources of its host to grow. These sugars are then used as building blocks and form the main structural component of the plant. Chlorophyll, a green-colored, magnesium-containing pigment is essential to this process; it is generally present in plant leaves, and often in other plant parts as well.

Plants usually rely on soil primarily for support and water (in quantitative terms), but also obtain compounds of nitrogen, phosphorus, and other crucial elemental nutrients. Epiphytic and lithophytic plants often depend on rainwater or other sources for nutrients and carnivorous plants supplement their nutrient requirements with insect prey that they capture. For the majority of plants to grow successfully they also require oxygen in the atmosphere and around their roots for respiration. However, some plants grow as submerged aquatics, using oxygen dissolved in the surrounding water, and a few specialized vascular plants, such as mangroves, can grow with their roots in anoxic conditions.

The leaf is usually the primary site of photosynthesis in plants.

Factors affecting growth

The genotype of a plant affects its growth, for example selected varieties of wheat grow rapidly, maturing within 110 days, whereas others, in the same environmental conditions, grow more slowly and mature within 155 days.[19]

Growth is also determined by environmental factors, such as temperature, available water, available light, and available nutrients in the soil. Any change in the availability of these external conditions will be reflected in the plants growth.

Biotic factors are also capable of affecting plant growth. Plants compete with other plants for space, water, light and nutrients. Plants can be so crowded that no single individual produces normal growth. Optimal plant growth can be hampered by grazing animals, suboptimal soil composition, lack of mycorrhizal fungi, and attacks by insects or plant diseases, including those caused by bacteria, fungi, viruses, and nematodes.[19]

Simple plants like algae may have short life spans as individuals, but their populations are commonly seasonal. Other plants may be organized according to their seasonal growth pattern: annual plants live and reproduce within one growing season, biennial plants live for two growing seasons and usually reproduce in second year, and perennial plants live for many growing seasons and continue to reproduce once they are mature. These designations often depend on climate and other environmental factors; plants that are annual in alpine or temperate regions can be biennial or perennial in warmer climates. Among the vascular plants, perennials include both evergreens that keep their leaves the entire year, and deciduous plants which lose their leaves for some part of it. In temperate and boreal climates, they generally lose their leaves during the winter; many tropical plants lose their leaves during the dry season.

The growth rate of plants is extremely variable. Some mosses grow less than 0.001 millimeters per hour (mm/h), while most trees grow 0.025-0.250 mm/h. Some climbing species, such as kudzu, which do not need to produce thick supportive tissue, may grow up to 12.5 mm/h.

Dried dead plants

Plants protect themselves from frost and dehydration stress with antifreeze proteins, heat-shock proteins and sugars (sucrose is common). LEA (Late Embryogenesis Abundant) protein expression is induced by stresses and protects other proteins from aggregation as a result of desiccation and freezing.[20]

Plant cell

Plant cell structure Main article: Plant cell

Plant cells are typically distinguished by their large water-filled central vacuole, chloroplasts, and rigid cell walls that are made up of cellulose, hemicellulose, and pectin. Cell division is also characterized by the development of a phragmoplast for the construction of a cell plate in the late stages of cytokinesis. Just as in animals, plant cells differentiate and develop into multiple cell types. Totipotent meristematic cells can differentiate into vascular, storage, protective (e.g. epidermal layer), or reproductive tissues, with more primitive plants lacking some tissue types.[21]

Physiology

Main article: Plant physiology

Photosynthesis

Main articles: Photosynthesis and Biological pigment

Plants are photosynthetic, which means that they manufacture their own food molecules using energy obtained from light. The primary mechanism plants have for capturing light energy is the pigment chlorophyll. All green plants contain two forms of chlorophyll, chlorophyll a and chlorophyll b. The latter of these pigments is not found in red or brown algae.

Immune system

By means of cells that behave like nerves, plants receive and distribute within their systems information about incident light intensity and quality. Incident light which stimulates a chemical reaction in one leaf, will cause a chain reaction of signals to the entire plant via a type of cell termed a "bundle sheath cell". Researchers from the Warsaw University of Life Sciences in Poland, found that plants have a specific memory for varying light conditions which prepares their immune systems against seasonal pathogens. [22]

Internal distribution

Main article: Vascular tissue

Vascular plants differ from other plants in that they transport nutrients between different parts through specialized structures, called xylem and phloem. They also have roots for taking up water and minerals. The xylem moves water and minerals from the root to the rest of the plant, and the phloem provides the roots with sugars and other nutrient produced by the leaves.[21]

Ecology

Main article: Plant ecology

The photosynthesis conducted by land plants and algae is the ultimate source of energy and organic material in nearly all ecosystems. Photosynthesis radically changed the composition of the early Earth's atmosphere, which as a result is now 21% oxygen. Animals and most other organisms are aerobic, relying on oxygen; those that do not are confined to relatively rare anaerobic environments. Plants are the primary producers in most terrestrial ecosystems and form the basis of the food web in those ecosystems. Many animals rely on plants for shelter as well as oxygen and food.

Land plants are key components of the water cycle and several other biogeochemical cycles. Some plants have coevolved with nitrogen fixing bacteria, making plants an important part of the nitrogen cycle. Plant roots play an essential role in soil development and prevention of soil erosion.

Distribution

This section requires expansion.

Plants are distributed worldwide in varying numbers. While they inhabit a multitude of biomes and ecoregions, few can be found beyond the tundras at the northernmost regions of continental shelves. At the southern extremes, plants have adapted tenaciously to the prevailing conditions. (See Antarctic flora.)

Plants are often the dominant physical and structural component of habitats where they occur. Many of the Earth's biomes are named for the type of vegetation because plants are the dominant organisms in those biomes, such as grasslands and forests.

Ecological relationships

The Venus flytrap, a species of carnivorous plant.

Numerous animals have coevolved with plants. Many animals pollinate flowers in exchange for food in the form of pollen or nectar. Many animals disperse seeds, often by eating fruit and passing the seeds in their feces. Myrmecophytes are plants that have coevolved with ants. The plant provides a home, and sometimes food, for the ants. In exchange, the ants defend the plant from herbivores and sometimes competing plants. Ant wastes provide organic fertilizer.

The majority of plant species have various kinds of fungi associated with their root systems in a kind of mutualistic symbiosis known as mycorrhiza. The fungi help the plants gain water and mineral nutrients from the soil, while the plant gives the fungi carbohydrates manufactured in photosynthesis. Some plants serve as homes for endophytic fungi that protect the plant from herbivores by producing toxins. The fungal endophyte, Neotyphodium coenophialum, in tall fescue (Festuca arundinacea) does tremendous economic damage to the cattle industry in the U.S.

Various forms of parasitism are also fairly common among plants, from the semi-parasitic mistletoe that merely takes some nutrients from its host, but still has photosynthetic leaves, to the fully parasitic broomrape and toothwort that acquire all their nutrients through connections to the roots of other plants, and so have no chlorophyll. Some plants, known as myco-heterotrophs, parasitize mycorrhizal fungi, and hence act as epiparasites on other plants.

Many plants are epiphytes, meaning they grow on other plants, usually trees, without parasitizing them. Epiphytes may indirectly harm their host plant by intercepting mineral nutrients and light that the host would otherwise receive. The weight of large numbers of epiphytes may break tree limbs. Hemiepiphytes like the strangler fig begin as epiphytes but eventually set their own roots and overpower and kill their host. Many orchids, bromeliads, ferns and mosses often grow as epiphytes. Bromeliad epiphytes accumulate water in leaf axils to form phytotelmata, complex aquatic food webs.[23]

Approximately 630 plants are carnivorous, such as the Venus Flytrap (Dionaea muscipula) and sundew (Drosera species). They trap small animals and digest them to obtain mineral nutrients, especially nitrogen and phosphorus.[24]

Importance

Potato plant. Potatoes spread to the rest of the world after European contact with the Americas in the late 1400s and early 1500s and have since become an important field crop. Timber in storage for later processing at a sawmill. A section of a Yew branch showing 27 annual growth rings, pale sapwood and dark heartwood, and pith (centre dark spot). The dark radial lines are longitudinal sections of small branches which became included by growth of the tree.

The study of plant uses by people is termed economic botany or ethnobotany; some consider economic botany to focus on modern cultivated plants, while ethnobotany focuses on indigenous plants cultivated and used by native peoples. Human cultivation of plants is part of agriculture, which is the basis of human civilization. Plant agriculture is subdivided into agronomy, horticulture and forestry.

Food

Much of human nutrition depends on land plants, either directly or indirectly. Human nutrition depends to a large extent on cereals, especially maize (or corn), wheat and rice. Other staple crops include potato, cassava, and legumes. Human food also includes vegetables, spices, and certain fruits, nuts, herbs, and edible flowers. Beverages produced from plants include coffee, tea, wine, beer and alcohol. Sugar is obtained mainly from sugar cane and sugar beet. Cooking oils and margarine come from maize, soybean, rapeseed, safflower, sunflower, olive and others. Food additives include gum arabic, guar gum, locust bean gum, starch and pectin. Livestock animals including cows, pigs, sheep, and goats are all herbivores; and feed primarily or entirely on cereal plants, particularly grasses.

Nonfood products

Wood is used for buildings, furniture, paper, cardboard, musical instruments and sports equipment. Cloth is often made from cotton, flax or synthetic fibers derived from cellulose, such as rayon and acetate. Renewable fuels from plants include firewood, peat and many other biofuels. Coal and petroleum are fossil fuels derived from plants. Medicines derived from plants include aspirin, taxol, morphine, quinine, reserpine, colchicine, digitalis and vincristine. There are hundreds of herbal supplements such as ginkgo, Echinacea, feverfew, and Saint John's wort. Pesticides derived from plants include nicotine, rotenone, strychnine and pyrethrins. Drugs obtained from plants include opium, cocaine and marijuana. Poisons from plants include ricin, hemlock and curare. Plants are the source of many natural products such as fibers, essential oils, dyes, pigments, waxes, tannins, latex, gums, resins, alkaloids, amber and cork. Products derived from plants include soaps, paints, shampoos, perfumes, cosmetics, turpentine, rubber, varnish, lubricants, linoleum, plastics, inks, chewing gum and hemp rope. Plants are also a primary source of basic chemicals for the industrial synthesis of a vast array of organic chemicals. These chemicals are used in a vast variety of studies and experiments.

Aesthetic uses

Thousands of plant species are cultivated for aesthetic purposes as well as to provide shade, modify temperatures, reduce wind, abate noise, provide privacy, and prevent soil erosion. People use cut flowers, dried flowers and houseplants indoors or in greenhouses. In outdoor gardens, lawn grasses, shade trees, ornamental trees, shrubs, vines, herbaceous perennials and bedding plants are used. Images of plants are often used in art, architecture, humor, language, and photography and on textiles, money, stamps, flags and coats of arms. Living plant art forms include topiary, bonsai, ikebana and espalier. Ornamental plants have sometimes changed the course of history, as in tulipomania. Plants are the basis of a multi-billion dollar per year tourism industry which includes travel to arboretums, botanical gardens, historic gardens, national parks, tulip festivals, rainforests, forests with colorful autumn leaves and the National Cherry Blossom Festival. Venus Flytrap, sensitive plant and resurrection plant are examples of plants sold as novelties.

Scientific and cultural uses

Tree rings are an important method of dating in archeology and serve as a record of past climates. Basic biological research has often been done with plants, such as the pea plants used to derive Gregor Mendel's laws of genetics. Space stations or space colonies may one day rely on plants for life support. Plants are used as national and state emblems, including state trees and state flowers. Ancient trees are revered and many are famous. Numerous world records are held by plants. Plants are often used as memorials, gifts and to mark special occasions such as births, deaths, weddings and holidays. Plants figure prominently in mythology, religion and literature. The field of ethnobotany studies plant use by indigenous cultures which helps to conserve endangered species as well as discover new medicinal plants. Gardening is the most popular leisure activity in the U.S. Working with plants or horticulture therapy is beneficial for rehabilitating people with disabilities. Certain plants contain psychotropic chemicals which are extracted and ingested, including tobacco, cannabis (marijuana), and opium.

Negative effects

Weeds are plants that grow where people do not want them. People have spread plants beyond their native ranges and some of these introduced plants become invasive, damaging existing ecosystems by displacing native species. Invasive plants cause billions of dollars in crop losses annually by displacing crop plants, they increase the cost of production and the use of chemical means to control them affects the environment.

Plants may cause harm to people and animals. Plants that produce windblown pollen invoke allergic reactions in people who suffer from hay fever. A wide variety of plants are poisonous to people and/or animals. Several plants cause skin irritations when touched, such as poison ivy. Certain plants contain psychotropic chemicals, which are extracted and ingested or smoked, including tobacco, cannabis (marijuana), cocaine and opium. Smoking causes damage to health or even death, while some drugs may also be harmful or fatal to people[25][26]. Both illegal and legal drugs derived from plants may have negative effects on the economy, affecting worker productivity and law enforcement costs.[27][28] Some plants cause allergic reactions in people and animals when ingested, while other plants cause food intolerances that negatively affect health.

See also

Plants portal

References

  1. ^ Haeckel G (1866). Generale Morphologie der Organismen. Berlin: Verlag von Georg Reimer. pp. vol.1: i–xxxii, 1–574, pls I–II; vol. 2: i–clx, 1–462, pls I–VIII.
  2. ^ "plant[2 - Definition from the Merriam-Webster Online Dictionary"]. http://www.merriam-webster.com/dictionary/plant%5B2%5D. Retrieved 2009-03-25.
  3. ^ "plant (life form) -- Britannica Online Encyclopedia". http://www.britannica.com/EBchecked/topic/463192/plant. Retrieved 2009-03-25.
  4. ^ Margulis, L. (1974). "Five-kingdom classification and the origin and evolution of cells". Evolutionary Biology 7: 45–78.
  5. ^ a b c d e Raven, Peter H., Ray F. Evert, & Susan E. Eichhorn, 2005. Biology of Plants, 7th edition. (New York: W. H. Freeman and Company). ISBN 0-7167-1007-2.
  6. ^ Bremer, K. (1985). "Summary of green plant phylogeny and classification". Cladistics 1: 369–385. doi:10.1111/j.1096-0031.1985.tb00434.x.
  7. ^ Mishler, Brent D.; S. P. Churchill (1985). "Transition to a land flora: phylogenetic relationships of the green algae and bryophytes". Cladistics 1: 305–328. doi:10.1111/j.1096-0031.1985.tb00431.x.
  8. ^ Mishler, Brent D.; Louise A. Lewis; Mark A. Buchheim; Karen S. Renzaglia; D. J. Garbary; Carl F. Delwiche; F. W. Zechman; T. S. Kantz; & Ron L. Chapman (1994). "Phylogenetic relationships of the "green algae" and "bryophytes"". Annals of the Missouri Botanical Garden (Annals of the Missouri Botanical Garden, Vol. 81, No. 3) 81 (3): 451–483. doi:10.2307/2399900. http://jstor.org/stable/2399900.
  9. ^ Van den Hoek, C., D. G. Mann, & H. M. Jahns, 1995. Algae: An Introduction to Phycology. pages 343, 350, 392, 413, 425, 439, & 448 (Cambridge: Cambridge University Press). ISBN 0-521-30419-9
  10. ^ Van den Hoek, C., D. G. Mann, & H. M. Jahns, 1995. Algae: An Introduction to Phycology. pages 457, 463, & 476. (Cambridge: Cambridge University Press). ISBN 0-521-30419-9
  11. ^ Crandall-Stotler, Barbara. & Stotler, Raymond E., 2000. "Morphology and classification of the Marchantiophyta". page 21 in A. Jonathan Shaw & Bernard Goffinet (Eds.), Bryophyte Biology. (Cambridge: Cambridge University Press). ISBN 0-521-66097-1
  12. ^ Schuster, Rudolf M., The Hepaticae and Anthocerotae of North America, volume VI, pages 712-713. (Chicago: Field Museum of Natural History, 1992). ISBN 0-914-86821-7.
  13. ^ Goffinet, Bernard; William R. Buck (2004). "Systematics of the Bryophyta (Mosses): From molecules to a revised classification". Monographs in Systematic Botany (Missouri Botanical Garden Press) 98: 205–239.
  14. ^ Gifford, Ernest M. & Adriance S. Foster, 1988. Morphology and Evolution of Vascular Plants, 3rd edition, page 358. (New York: W. H. Freeman and Company). ISBN 0-7167-1946-0.
  15. ^ Taylor, Thomas N. & Edith L. Taylor, 1993. The Biology and Evolution of Fossil Plants, page 636. (New Jersey: Prentice-Hall). ISBN 0-13-651589-4.
  16. ^ International Union for Conservation of Nature and Natural Resources, 2006. IUCN Red List of Threatened Species:Summary Statistics
  17. ^ Kenrick, Paul & Peter R. Crane. 1997. The Origin and Early Diversification of Land Plants: A Cladistic Study. (Washington, D.C.: Smithsonian Institution Press). ISBN 1-56098-730-8.
  18. ^ Smith, Alan R., Kathleen M. Pryer, E. Schuettpelz, P. Korall, H. Schneider, & Paul G. Wolf. (2006). "A classification for extant ferns". Taxon 55(3): 705-731.
  19. ^ a b Robbins, W.W., Weier, T.E., et al., Botany:Plant Science, 3rd edition , Wiley International, New York, 1965.
  20. ^ Goyal, K., Walton, L. J., & Tunnacliffe, A. (2005). "LEA proteins prevent protein aggregation due to water stress". Biochemical Journal 388 (Part 1): 151 – 157. doi:10.1042/BJ20041931. PMID 15631617. PMC 1186703. Archived from the original on 2009-08-03. http://www.webcitation.org/5il9QhYT0.
  21. ^ a b Campbell, Reece, Biology, 7th edition, Pearson/Benjamin Cummings, 2005.
  22. ^ BBC Report
  23. ^ Howard Frank, Bromeliad Phytotelmata, October 2000
  24. ^ Barthlott, W., S. Porembski, R. Seine, and I. Theisen. 2007. The Curious World of Carnivorous Plants: A Comprehensive Guide to Their Biology and Cultivation. Timber Press: Portland, Oregon.
  25. ^ "cocaine/crack". http://www.urban75.com/Drugs/drugcoke.html.
  26. ^ "Deaths related to cocaine". http://ar2005.emcdda.europa.eu/en/page050-en.html.
  27. ^ "Illegal drugs drain $160 billion a year from American economy". Archived from the original on 2008-02-15. http://web.archive.org/web/20080215071055/http://www.whitehousedrugpolicy.gov/NEWS/press02/012302.html.
  28. ^ "The social cost of illegal drug consumption in Spain". http://www.ingentaconnect.com/content/bsc/add/2002/00000097/00000009/art00012.

Further reading

General
Species estimates and counts

External links

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Botany
Subdisciplines of botany Ethnobotany · Paleobotany · Plant anatomy · Plant ecology · Plant evo-devo · Plant morphology · Plant physiology
Plants Evolutionary history of plants · Algae · Bryophyte · Pteridophyte · Gymnosperm · Angiosperm
Plant parts Flower · Fruit · Leaf · Meristem · Root · Stem · Stoma · Vascular tissue · Wood
Plant cells Cell wall · Chlorophyll · Chloroplast · Photosynthesis · Plant hormone · Plastid · Transpiration
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Classification of Archaeplastida / Plantae sensu lato
Rhodophyta Rhodophyceae · Bangiophyceae · Florideophyceae · Goniotrichales · Stylonematophyceae
Glaucocystophyceae Glaucocystis · Cyanophora · Gloeochaete
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Prasinophyceae

UTC clade: Ulvophyceae · Trebouxiophyceae · Chlorophyceae
Streptophyta
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Chlorokybophyceae Chlorokybales
Klebsormidiophyceae Klebsormidiales
Zygnematophyceae Desmidiales · Zygnematales
Charophyceae Charales
Coleochaetophyceae Coleochaetales
Embryophyta/ Plantae sensu strictissimo
Bryophytes (non-vascular) Marchantiophyta · Anthocerotophyta · Bryophyta "Moss" · Horneophytopsida
Tracheophyta
Lycopodiophyta Isoetopsida (Isoetales, Selaginellales) · Lycopodiopsida (Lycopodiales)
Euphyllophyta Moniliformopses (Equisetopsida, Filicopsida, Psilotopsida) Spermatophyta: Gymnosperm (Pinophyta, Cycadophyta, Ginkgophyta, Gnetophyta) · Magnoliophyta
See also: list of plant orders
Eukaryota classification
Domain : Archaea · Bacteria · Eukaryota
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Archaeplastida, or Plantae sensu lato Viridiplantae/Plantae sensu stricto · Rhodophyta · Glaucocystophyceae
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How is plant life related to animal life? And Did animal life evolve from plant life?
Q. Also I'd like to know things like, Which came first, plant or animal life? (or did they come about at the same rate?) Are organisms like bacteria more closely related to plant or animal life? How did plant life begin? Intuitively, we sense there is something different about plant life from animal life - but what exactly is that difference? There are plants that can move quickly, like a Venus Flytrap, and animals that act more the way we think plants would act, like a sea anemone. Thanks for your answers!
Asked by Heron By The Sea - Sat Apr 12 16:02:33 2008 - - 1 Answers - 1 Comments

A. Animal life did not evolve from plant life. Both plant life and animal life evolved on parallel paths from common ancestors. That is why you see the overlap in structures that you mentioned. At one time there were only single celled life forms. At that point it would have been difficult to call them either plants or animals. From there several different lines evolved specializing in different ways which eventualy lead to all the plants and animals we know today. Some biologists have even said there are not clear distinctions between plants and animals, but that plants are really nothing but very slow moving animals.
Answered by polybisep - Sun Apr 13 03:26:09 2008

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